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After the frontier incident, everyone who knew the circumstances believed the Noise were gone and the pain they caused was at an end. But a new conflict approached unseen. In the Yokohama Harbor Oosan-bashi Pier, a new pattern is detected that is similar to the Noise. A combat group wielding mysterious technology stands in the way of Hibiki and the others. When she hears of this enemy of unprecedented strength, Tsubasa hurries back home from England. But the Symphogear users see no opening to counterattack are forced into a difficult fight. In this situation of extreme disparity, the battle for the song that will end the world begins.
Peace had seemed to return to the world when Hibiki, Tsubasa, and Chris had saved the planet from the Luna Attack with their song. However, 3 months later at a joint concert between Tsubasa Kazanari and foreign superstar Maria Cadenzavna Eve, the Symphogear team is faced with her dangerous declaration of war against the world. Maria and her allies Kirika and Shirabe have a fearful power backing them: The power of a dark Symphogear.
Legendary rap group The Dirty Dawg could have taken Japan by storm, but they broke up before they could. Now each member has joined one of four rival groups—fighting for the territories Ikebukuro, Yokohama, Shibuya, and Shinjuku. Battles for turf are fought with Hypnosis Mics. These special microphones have the power to affect the human spirit, and people now use rap to determine superiority.
One day a black meteorite fell on the world, and the world completely changed. The meteorite produced grotesque monsters called D2, which started running rampant. D2 very quickly banned all music, which was the one thing able to overcome them. But there were some people who resisted the D2. They are young women who hold the power of music, the "Musicart." These young women hold "scores" that are able to defeat the monsters. And there are also people who lead these women, the Conductor.