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The Muromachi Shogunate is weakening due to an extended war and the country has entered the tumultuous time known as the Warring States Period where blood is washed away by more blood. In the depths of the deep forests in the unexplored Kinokuni area, the Saika Ikki mercenaries are armed with guns and waving the Yatagarasu flag. Their leader is a foreigner named Magoichi. Meanwhile, a man named Saburou struggles to protect Hinomoto from European invasion. The fate of thes two will lead them both to cross swords with invaders.
It's 1923, and vampires have existed for quite a while. But now, the number of vampires is increasing. and a mysterious, artificial blood source called Ascra has appeared. The Japanese government, in turn, creates “Code Zero,” a unit within the army tasked with taking down the vampiric forces. And what better way to track vampires than by using vampires? Created by Lieutenant General Nakajima, this unit has historically been in the business of information war, but has been re-assigned to solve the vampire crisis. It's up to Code Zero and the S-class vampire Deffrot to investigate this increase and put a stop to it before society crumbles.
This is the story of the extraordinary Benkei, who after having met his enemy Yoshitsune in his childhood (then named 'Ushiwaka') came to guard him as a lifetime partner in later years.