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Once there lived an eccentric author called Drosselmeyer who wrote grand tragedies - one of them was the tale of a prince who sealed away an evil raven by breaking his own heart into tiny pieces. However, before the story could be completed, the author died and the tale took on a life of its own. Now, in a town where fiction and reality meet, the story continues on its tragic course with Ahiru, a duck who transforms into the beautiful Princess Tutu in order to restore the prince's heart. But will Ahiru's act of love be enough to defy the story's terrible destiny and lead to a happy ending?